1. Institute of Zoology; Zoological Society of London; Regent's Park London NW1 4RY UK
2. Department of Genetics, Evolution and Environment; University College London; Gower Street London WC1E 6BT UK
3. School of Biological Sciences; University of Adelaide; Adelaide SA 5005 Australia
4. Centre for Invasion Biology; Department of Botany and Zoology; Stellenbosch University; Stellenbosch South Africa