1. Department of Molecular Microbiology; Washington University School of Medicine; St Louis MO 63105 USA
2. Institute for Cell and Molecular Biosciences; The Medical School; Newcastle University; Newcastle upon Tyne NE2 4HH UK
3. Howard Hughes Medical Institute; New Haven CT 06536-0812 USA
4. Department of Microbial Pathogenesis; Yale School of Medicine; New Haven CT 06536 USA
5. Yale Microbial Diversity Institute; West Haven CT 06516 USA