1. Plant Germplasm and Genomics Center, Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Department of Ecology and Evolutionary Biology University of Toronto Toronto Ontario M5S 3B2 Canada
4. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China