1. School of Life Sciences University of Warwick Wellesbourne Campus Warwickshire CV35 9EF UK
2. Functional and Comparative Genomics Institute of Integrative Biology University of Liverpool Liverpool L69 7ZB UK
3. Applied Statistical Solutions Bishops Tachbrook Leamington CV33 9RJ UK
4. Department of Crop Genetics John Innes Centre Norwich Research Park Norwich NR4 7UH UK