1. Key Laboratory of Freshwater Fisheries and Germplasm Resource Utilization; Freshwater Fisheries Research Center; Chinese Academy of Fishery Sciences; Ministry of Agriculture; Wuxi 214081 Jiangsu China
2. Wuxi Fisheries College; Nanjing Agricultural University; Wuxi 214081 Jiangsu China
3. School of Life Sciences; Huaiyin Normal University; Huaian 223300 Jiangsu China