1. Department of Biological Sciences; National University of Singapore; 14 Science Drive 4 Singapore 117543 Singapore
2. Centre for Tropical Biodiversity and Climate Change; School of Marine and Tropical Biology; James Cook University; Townsville QL 4811 Australia
3. Centre for Tropical Environmental and Sustainability Science (TESS) and School of Marine and Tropical Biology; James Cook University; Cairns QL 4878 Australia
4. Department of Animal and Plant Sciences; University of Sheffield; Western Bank Sheffield S10 2TN UK
5. Herpetology Section, Zoology Division; National Museum of the Philippines; Padre Burgos Avenue Ermita 1000 Manila Philippines