1. NIOZ Royal Netherlands Institute for Sea Research; Department of Coastal Systems and Utrecht University; Den Burg, Texel 1790 AB The Netherlands
2. Department of Vertebrate Zoology; Lomonosov Moscow State University; 119991 Moscow Russia
3. Conservation Ecology Group; Groningen Institute for Evolutionary Life Sciences (GELIFES); University of Groningen; 9700 CC Groningen The Netherlands