1. Department of Ecology and Evolutionary Biology; University of Michigan; Ann Arbor MI 48109 USA
2. NOAA Great Lakes Environmental Research Laboratory; Ann Arbor MI 48108 USA
3. Department of Earth and Environmental Sciences; University of Michigan; Ann Arbor MI 48109 USA
4. Cooperative Institute for Limnology and Ecosystems Research; University of Michigan; Ann Arbor MI 48109 USA