1. State Key Laboratory for Agrobiotechnology, Frontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), and Beijing Key Laboratory of Crop Genetic Improvement China Agricultural University Beijing 100193 China
2. Council for Agricultural Research and Economics, Research Centre for Cereal and Industrial Crops I‐24126 Bergamo Italy
3. National Key Laboratory of Crop Genetic Improvement and National Center of Plant Gene Research (Wuhan) Huazhong Agricultural University Wuhan 430070 China