1. Department of Biology; Laboratory of Functional Morphology; University of Antwerp; Universiteitsplein 1 2610 Wilrijk Belgium
2. Centre for Invasion Biology; Stellenbosch University; Private Bag X1 Matieland 7602 South Africa
3. Applied Biodiversity Research Division; South African National Biodiversity Institute; Private Bag X7 Claremont 7735 Cape Town South Africa