1. Department of Botany and Zoology; Stellenbosch University, Private Bag X1, Matieland; 7602 Stellenbosch South Africa
2. Prizker Laboratory for Molecular Systematics and Evolution, The Field Museum, 1400 South Lake Shore Drive; Chicago IL 60605 U.S.A.
3. Museum of Vertebrate Zoology and Department of Integrative Biology; University of California, Berkeley, 3101 Valley Life Sciences Building; Berkeley CA 94720 U.S.A.