1. Department of Biochemistry and Molecular Biology; Faculty of Agriculture and Life Science; Hirosaki University; Hirosaki Aomori Japan
2. Division of Morphogenesis; National Institute for Basic Biology; National Institutes of Natural Sciences; Okazaki Aichi Japan
3. Department of Basic Biology; School of Life Science; the Graduate University of Advanced Studies; Hayama Kanagawa Japan
4. Department of Developmental Biology and Neurosciences; Graduate School of Life Sciences; Tohoku University; Sendai Japan