1. The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, School of Life Science Shandong University Qingdao 266237 China
2. Grassland Agri‐Husbandry Research Center, College of Grassland Science Qingdao Agricultural University Qingdao 266109 China
3. Institute for Agricultural Biosciences Oklahoma State University 3210 Sam Noble Parkway Ardmore OK 73401 USA