1. Atmosphere and Ocean Research Institute University of Tokyo 5‐1‐5, Kashiwanoha Kashiwa Chiba 277‐8564 Japan
2. Fisheries Research Agency National Research Institute of Fisheries Science Komaki 1088 Ueda Nagano 386‐0031 Japan
3. Department of Biology Osaka Medical College Daigaku‐machi 2‐7 Takatsuki Osaka 569‐8686 Japan