1. Department of Botany; Smithsonian Institution; NMNH MRC-166, P.O. Box 37012 Washington DC 20013
2. Department of Organismic and Evolutionary Biology; Harvard University Herbaria; 22 Divinity Avenue Cambridge Massachusetts 02138
3. Department of Ecology and Evolutionary Biology and University of Michigan Herbarium; 3600 Varsity Drive Ann Arbor Michigan 48108
4. Real Jardín Botánico; RJB-CSIC; Plaza de Murillo 2 28014 Madrid Spain
5. The Huntington Botanical Gardens; 1151 Oxford Road San Marino California 91108