1. State Key Laboratory for Quality and Safety of Agro‐products, Institute of Plant Virology Ningbo University Ningbo China
2. College of Plant Protection Northwest Agriculture and Forestry University Yangling China
3. Institute of Crop Sciences, State Key Laboratory of Crop Gene Resources and Breeding, National Key Facility for Crop Gene Resources and Genetic Improvement, Chinese Academy of Agricultural Sciences Beijing China
4. School of Life Sciences University of Dundee Dundee UK