1. State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, Hubei, 430072, PR China
2. University of Chinese Academy of Sciences, Beijing, 100049, PR China
3. Department of Wildlife, Fisheries, and Aquaculture, Mississippi State UniversityMississippi State, MS 39762, USA
4. School of Life Sciences, Jianghan University, Wuhan, Hubei, 430058, PR China