1. Department of Biology, Institute for Great Lakes Research, Central Michigan University, Biosciences 2100, 1455 Calumet Court, Mount Pleasant, MI 48859, USA.
2. Great Lakes Laboratory for Fisheries and Aquatic Sciences, Fisheries and Oceans Canada, 867 Lakeshore Road, Burlington, ON L7S 1A1, Canada.
3. Ontario Ministry of Natural Resources and Forestry, 300 Water Street, Peterborough, ON K9J 8M5, Canada.