1. Center for Watershed Sciences/California Trout, University of California, One Shields Avenue, Davis, CA 95616, USA.
2. Department of Wildlife, Fish, and Conservation Biology, University of California, One Shields Ave., Davis, CA 95616, USA.
3. Aquatic Health Program, School of Veterinary Medicine, Department of Anatomy, Physiology, and Cell Biology, University of California, One Shields Ave., Davis, CA 95616, USA.
4. Center for Watershed Sciences, University of California, One Shields Ave., Davis, CA 95616, USA.
5. Fisheries Ecology Division, Southwest Fisheries Science Center, National Marine Fisheries Service, National Oceanic and Atmospheric Administration, 110 McAllister Way, Santa Cruz, CA 95060, USA.
6. Institute of Marine Sciences, University of California, 1156 High Street, Santa Cruz, CA 95064, USA.