1. Estuary Research Center, Shimane University, Matsue, Shimane, Japan.
2. Demersal Fish Resources Division, Fisheries Stock Assessment Center, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, Niigata, Niigata, Japan.
3. Socio-Ecological Systems Division, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, Yokohama, Kanagawa, Japan.
4. Graduate School of Environmental Science, Hokkaido University, Hakodate, Japan.
5. Faculty of Fisheries Science, Hokkaido University, Hakodate, Hokkaido, Japan.
6. Highly Migratory Resources Division, Fisheries Stock Assessment Center, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, Yokohama, Kanagawa, Japan.
7. Fisheries and Oceans Hakodate, Hakodate, Hokkaido, Japan.
8. Field Science Center for Northern Biosphere, Hokkaido University, Hakodate, Hokkaido, Japan.