1. Fisheries and Oceans Canada, Cooperative Resource Management Institute, School of Resource and Environmental Management, Simon Fraser University, Burnaby, BC V5A 1S6, Canada
2. Earth to Ocean Research Group, Simon Fraser University, Burnaby, BC V5A 1S6, Canada
3. Fisheries and Oceans Canada, Cultus Lake Salmon Research Laboratory, Cultus Lake, BC V2R 5B6, Canada