1. Department of Forest Resources Management, Faculty of Forestry, University of British Columbia, 2424 Main Mall, Vancouver, BC V6T 1Z4, Canada.
2. Department of Forest Sciences and Centre for Applied Conservation Research, Faculty of Forestry, University of British Columbia, 2424 Main Mall, Vancouver, BC V6T 1Z4, Canada.
3. British Columbia Ministry of Forests and Range, 2100 Labieux Road, Nanaimo, BC V9T 6E9, Canada.