1. School of the Environment, Washington State University, P.O. Box 642812, Pullman, WA 99164, USA.
2. Department of Biology, Trent University, Peterborough, ON K9J 0G2, Canada.
3. Washington State Department of Natural Resources, Northeast Region, 225 S. Silke Road, Colville, WA 99114, USA.
4. Washington State Department of Fish and Wildlife, Okanogan District, 350 Bear Creek Road, Winthrop, WA 98862, USA.
5. Okanogan-Wenatchee National Forest, 24 West Chewuch, Winthrop, WA 98862, USA.