1. School of Ocean Sciences, College of Natural Sciences, Bangor University, Menai Bridge, Anglesey, LL59 5AB, United Kingdom.
2. KwaZulu-Natal Sharks Board, Private Bag 2, Umhlanga Rocks 4320, South Africa.
3. Biomedical Research Unit, University of KwaZulu-Natal, P.O. Box X54001, Durban 4000, South Africa.
4. Great Lakes Institute of Environmental Research, University of Windsor, Windsor, ON N9B 3P4, Canada.