1. Department of Renewable Resources, University of Alberta, 751 General Services Building, Edmonton, AB T6G 2H1, Canada.
2. Department of Forest Resources Management, University of Alberta, 751 General Services Building, Edmonton, AB T6G 2H1, Canada.
3. Département des sciences du bois et de la forêt, Faculté de foresterie, de géographie et de géomatique, Université Laval, 2425, rue de la Terrasse, Pavillon Gene-H.-Kruger, Québec, QC G1V 0A6, Canada.