1. Royal Alberta Museum, 9810 103a Avenue NW, Edmonton, AB T5J 0G2, Canada.
2. Alberta Biodiversity Monitoring Institute, CW 405, Biological Sciences Building, University of Alberta, Edmonton, AB T6G 2E9, Canada.
3. Department of Renewable Resources, Faculty of Agricultural, Life & Environmental Sciences, University of Alberta, Edmonton, AB T6G 2E9, Canada.
4. Eastern Oregon Agricultural Research Center, United States Department of Agriculture, 67826-A Hwy 205, Burns, OR 97720, USA.
5. Sierra Foothills Research and Extension Center, University of California Division of Agriculture and Natural Resources, Browns Valley, CA 95918, USA.