1. School of Environmental Design and Rural Development, University of Guelph, 50 Stone Rd E, Guelph, ON, N1G 2W1, Canada
2. Ikaarvik Barriers to Bridges, Hamlet of Pond Inlet, Nunavut, X0A 0S0, Canada
3. Wildlife Research Division, Environment and Climate Change Canada, 105 McGill Street, Montreal, QC H2Y 2E7, Canada