1. Department of Biology, Queen’s University, 116 Barrie Street, Kingston, ON K7L 3N6, Canada.
2. Department of Biological Sciences, Thompson Rivers University, 805 TRU Way, Kamloops, BC V2C 0C8, Canada.
3. Canadian Wildlife Service, Environment and Climate Change Canada, 115 Perimeter Road, Saskatoon, SK, S7N 0X4, Canada.