1. Jiaozhou Bay National Marine Ecosystem Research Station, Institute of Oceanology, Chinese Academy of Sciences, 7 Nanhai Road, Qingdao 266071, China.
2. University of Chinese Academy of Sciences, Beijing 100049, China.
3. Laboratory for Marine Ecology and Environmental Science, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266071, China.
4. Center for Ocean Mega Science, Chinese Academy of Sciences, Qingdao 266071, China.