1. Department of Veterinary Medicine, Faculty of Agriculture, Iwate University, Morioka 020–8550, Japan
2. The Iwate Research Center for Wildlife Diseases, Morioka 020–0816, Japan
3. The Sarudokonet, NPO for Wildlife Management, Matsusaka 519–2144, Japan
4. Cryobiofrontier Research Center, Faculty of Agriculture, Iwate University, Morioka 020–8550, Japan