1. Laboratory of Wildlife Biology and Medicine, Faculty of Veterinary Medicine, Hokkaido University, Hokkaido, Japan
2. Laboratory of Clinical Veterinary Medicine for Large Animal, School of Veterinary Medicine, Kitasato University, Aomori, Japan
3. Laboratory of Theriogenology, Faculty of Veterinary Medicine, Hokkaido University, Hokkaido, Japan
4. Laboratory of Animal Reproduction, School of Veterinary Medicine, Kitasato University, Aomori, Japan
5. Primate Research Institute, Kyoto University, Aichi, Japan