1. Department of Biology, Tennessee Technological University, 1100 North Dixie Avenue, Cookeville, TN 38505, USA
2. Department of Biology, Framingham State University, 100 State Street, Framingham, MA 01701, USA
3. The Nature Conservancy, 701 West Ocean Acres Drive, Kill Devil Hills, NC 27948, USA
4. Department of Biological Sciences, University of Cincinnati, Rieveschl Hall, 318 College Drive, Cincinnati, OH 45221, USA