1. Department of Biology, Florida Southern College, Lakeland, FL 33801, USA
2. Department of Biology, Alma College, Alma, MI 48801, USA
3. Department of Mathematics, Alma College, Alma, MI 48801, USA
4. Biology Department, Grand Valley State University, Allendale, MI 49401, USA
5. Colegio de Ciencias Biológicas y Ambientales COCIBA & Galápagos Science Center GSC, Universidad San Francisco de Quito, Ecuador