1. Department of Ecology, Evolution, and Behavior, University of Minnesota, 1987 Upper Buford Circle, Saint Paul, MN 55108, USA
2. Institute for Northern Studies, University of the Highlands and Islands, Scott's House, Grainshore Road, Hatston, Orkney, KW15 1FL, UK
3. Bell Museum and Department of Fisheries, Wildlife, and Conservation Biology, University of Minnesota, 1987 Upper Buford Circle, Saint Paul, MN 55108, USA