1. US Geological Survey, Fort Collins Science Center, 2150 Centre Avenue, Building C, Fort Collins, CO 80526, USA
2. US Geological Survey, Northern Rocky Mountain Science Center, Wildlife Biology Program, University of Montana, Missoula, MT 59812, USA
3. US Geological Survey, Patuxent Wildlife Research Center, SO Conte Anadromous Fish Laboratory, One Migratory Way, Turners Falls, MA 01376, USA
4. US Geological Survey, Forest and Rangeland Ecosystem Science Center, 970 Lusk Street, Boise, ID 83706, USA
5. University of Vermont, Rubenstein School of Environment and Natural Resources, Aiken Center, 81 Carrigan Drive, Burlington, VT 05405, USA