1. Cornell Wildlife Health Laboratory, Animal Health Diagnostic Center, Cornell School of Veterinary Medicine, 240 Farrier Road, Ithaca, New York 14853, USA
2. Department of Population Medicine and Diagnostic Sciences, Animal Health Diagnostic Center, Cornell School of Veterinary Medicine, 240 Farrier Road, Ithaca, New York 14853, USA
3. Department of Biomedical Sciences, Cornell School of Veterinary Medicine, 240 Farrier Road, Ithaca, New York 14853, USA
4. Wildlife Health Unit, New York State Department of Environmental Conservation, 108 Game Farm Road, Delmar, New York 12054, USA
5. Virology Department, Animal and Plant Health Agency-Weybridge, New Haw, Addlestone, Surrey, KT15 3NB, UK