1. US Geological Survey, National Wildlife Health Center, Honolulu Field Station, PO Box 50167, Honolulu, Hawaii 96850, USA
2. US Fish & Wildlife Service, Midway Atoll National Wildlife Refuge, PO Box 50167, Honolulu, Hawaii 96850, USA
3. US Geological Survey, Pacific Islands Ecosystem Science Center, PO Box 44, Hawaii National Park, Hawaii 96718, USA
4. US Geological Survey, National Wildlife Health Center, 6006 Schroeder Rd., Madison, Wisconsin 53711, USA