1. Department of Wildlife and Animal Resource Management, Makerere University, Kampala, Uganda
2. Morris Animal Foundation's Mountain Gorilla Veterinary Project, Baltimore, Maryland 21217, USA
3. Department of Parasitology and Microbiology, Makerere University, Kampala, Uganda
4. Uganda Wildlife Authority, Kampala, Uganda
5. Medical Department, The Baltimore Zoo, Baltimore, Maryland 21217, USA
6. Division of Comparative Medicine, School of Medicine, Johns Hopkins University, Baltimore, Maryland 21205, USA
7. Department of Molecular Microbiology and Immunology, and Department of Environmental Health Sciences, School of Hygiene and Public Health, Johns Hopkins University, Baltimore, Maryland 21205, USA