1. Institute for Evolutionary Ecology and Conservation Genomics, Albert-Einstein Allee 11, 89081 Ulm, Germany
2. Large Animal Research Group, Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, United Kingdom
3. University of Pretoria, Mammal Research Institute, Private Bag X20, Hatfield 0028, Pretoria, South Africa