1. Laboratory of Food Microbiology and Food Safety, School of Veterinary Medicine, Rakuno Gakuen University, Ebetsu 069-8501, Japan
2. Miyajimanuma Waterbird and Wetland Center, Oomagari 3, Nishibibai, Bibai 072-0057, Japan
3. Division of Bioresources, Hokkaido University Research Center for Zoonosis Control, Kita 20, Nishi 10, Kita-ku, Sapporo 001-0020, Japan
4. Faculty of Global Human Sciences, Kobe University, 3-11, Tsurukabuto, Nada-ku, Kobe, Hyogo 657-8501, Japan
5. Graduate School of Human Development and Environment, Kobe University, 3-11, Tsurukabuto, Nada-ku, Kobe, Hyogo 657-8501, Japan
6. Laboratory of Environmental Geochemistry, College of Agriculture, Food and Environment Sciences, Rakuno Gakuen University, Ebetsu 069-8501, Japan
7. Department of Applied Veterinary Science, The United Graduated School of Veterinary Science, Gifu University, Yanagido Post 501-1193, Japan