1. Odum School of Ecology, University of Georgia, 140 E. Green Street, Athens, Georgia, 30602, USA
2. These authors contributed equally
3. Current address: School of Forestry and Wildlife Sciences, Auburn University, 3301 Forestry and Wildlife Sciences Building, 602 Duncan Drive, Auburn, Alabama 36849, USA
4. Current address: School of Biological Sciences, University of Texas at Austin, 100 E. 24th Street, NHB 2.606, Austin, Texas 78712, USA
5. Southeastern Cooperative Wildlife Disease Study, College of Veterinary Medicine, University of Georgia, 589 D. W. Brooks Drive, Athens, Georgia 30602, USA
6. Department of Infectious Diseases, College of Veterinary Medicine, University of Georgia, 501 D. W. Brooks Drive, Athens, Georgia 30602, USA