1. School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia 5005, Australia
2. Current address: School of Earth and Environmental Sciences, University of Adelaide, Adelaide, South Australia 5005, Australia
3. School of Medical Sciences, University of Adelaide, Adelaide, South Australia 5005, Australia
4. Veterinary Department, Royal Zoological Society of South Australia, Frome Road, Adelaide, South Australia 5000, Australia