1. Department of Biosciences, Nottingham Trent University, Nottingham NG1 4FQ, UK
2. Present address: Berkshire and Surrey Pathology Services, Frimley Health NHS Trust, Wexham Park Hospital, Slough, UK
3. Life Sciences, University of Westminster, London, UK
4. Department of Food and Nutritional Sciences, University of Reading, Reading, UK
5. SRUC Veterinary Services, Inverness, UK