1. Centro Nacional de Microbiología, Instituto de Salud Carlos III, Madrid 28220, Spain
2. Centro de Investigación Biomédica de Epidemiología y Salud Pública, CIBERESP, Spain
3. Wildlife Zoonoses and Vector-borne Diseases Research Group (OIE Reference Laboratory/WHO Collaborating Centre), Animal Health and Veterinary Laboratories Agency (AHVLA, Weybridge), New Haw, Addlestone, Surrey KT15 3NB, UK
4. Estación Biológica de Doñana, CSIC, Seville 41092, Andalusia, Spain
5. Department of Zoology and Animal Cell Biology, University of the Basque Country (UPV/EHU), Leioa 48940, The Basque Country, Spain
6. University of Liverpool, National Consortium for Zoonosis Research, Leahurst, Neston, South Wirral CH64 7TE, UK