1. School of Food & Biological Engineering, Jiangsu University, Zhenjiang 212013, PR China
2. Second Institute of Oceanography, State Oceanic Administration, Hangzhou 310012, PR China
3. Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, PR China
4. Institute of Life Sciences, and the Moshe Shilo Minerva Center for Marine Biogeochemistry, The Hebrew University of Jerusalem, Jerusalem 91904, Israel