1. Department of Veterinary Preventive Medicine, College of Veterinary Medicine, Jilin University, Xi'an Road 5333, Changchun, Jilin 130062, PR China
2. College of Quartermaster Technology Science, Jilin University, Xi'an Road 5333, Changchun, Jilin 130062, PR China
3. CAS Key Laboratory of Pathogenic Microbiology and Immunology, Institute of Microbiology, Chinese Academy of Sciences, Beijing, PR China
4. China Animal Health and Epidemiology Center, Qingdao, Nanjing Road 369, Qingdao, Shandong 266032, PR China
5. Comprehensive Pneumology Center, Institute of Lung Biology and Disease (iLBD), Helmholtz Zentrum München, Ingolstädter Landstraße 1, D-85764 Neuherberg/Munich, Germany