1. Univ Brest, CNRS, Ifremer, Unité Biologie et Ecologie des Ecosystèmes marins Profonds BEEP, UMR 6197, IRP 1211 MicrobSea, IUEM, Rue Dumont d’Urville, F-29280 Plouzané, France
2. Key Laboratory of Marine Genetic Resources, Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, PR China