1. Japan International Research Center for Agricultural Sciences, Tsukuba, Ibaraki 305-8686, Japan
2. Hainan University, Haikou, Hainan 570228, PR China
3. Zhengzhou University, Zhengzhou, Henan 450001, PR China
4. Japan Collection of Microorganisms, RIKEN BioResource Center, Wako, Saitama 351-0198, Japan