1. Division of Biology, Faculty of Sciences, Rajamangala University of Technology Thanyaburi, Pathumthani 33130, Thailand
2. Department of Microbiology, Faculty of Sciences, Chulalongkorn University, Bangkok 10330, Thailand
3. Japan Collection of Microorganisms, RIKEN BioResource Center, 3-1-1 Koyadai, Tsukuba, Ibaraki 305-0074, Japan
4. Department of Biochemistry and Microbiology, Faculty of Pharmaceutical Sciences, Chulalongkorn University, Bangkok 10330, Thailand