1. Biotechnology Research Division, National Institute of Fisheries Science (NIFS), Gijang, Busan 46083, Republic of Korea
2. Jeju Fisheries Research Institute, National Institute of Fisheries Science (NIFS), Jeju 63610, Republic of Korea
3. Department of Food Science and Biotechnology, Sungkyunkwan University, Jangan-gu, Suwon 16419, Republic of Korea